1.2 ceph http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Young, Richard E., Michael Vecchione, and Katharina M. Mangold (1922-2003). 2012. Cephalopoda Cuvier 1797. Octopods, squids, nautiluses, etc.. Version 10 November 2012 (under construction). http://tolweb.org/Cephalopoda/19386/2012.11.10 in The Tree of Life Web Project, http://tolweb.org/ is about todo - mireot definition plural term taxonomic disambiguation term used in octopoda term used in squid source synonym_type_property has_alternative_id has_broad_synonym database_cross_reference has_exact_synonym has_narrow_synonym has_obo_format_version has_related_synonym has_synonym_type in_subset shorthand obo-source part_of BFO:0000050 part_of has part BFO:0000051 has_part homologous_to develops from capable_of adjacent to aboral to Away from or opposite the mouth. Away from or opposite the mouth. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 continuous_with dorsal_to proximal_to secretes surrounded_by surrounds trait of ventral_to depicts eyelid 2011-01-30T08:15:10Z Consider re-using Uberon class cjm accessory nidamental gland The basic structural unit is a tubule composed of a single layer of epithelial cells containing ordered arrays of rough endoplasmic reticulum and a lumenal surface covered with microvilli, cilia, and structural specialization presumed to be involved in secretion. The basic structural unit is a tubule composed of a single layer of epithelial cells containing ordered arrays of rough endoplasmic reticulum and a lumenal surface covered with microvilli, cilia, and structural specialization presumed to be involved in secretion. cephalopod eye 2011-01-30T08:21:37Z cjm todo - reconcile with camera-type eye in uberon circumoral appendage bud accessory gland complex The glandular complex in cirrates that forms sperm packets and is a counterpart of the spermatophore-forming complex of other cephalopods. The glandular complex in cirrates that forms sperm packets and is a counterpart of the spermatophore-forming complex of other cephalopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 acetabulum A sucker of the sepia or cuttlefish and related animals. A sucker of the sepia or cuttlefish and related animals. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 adult stage A female that has mature eggs (these are frequently stored in the oviducts), or a male that has produced spermatophores (these are stored in Needham's sac). A female that has mature eggs (these are frequently stored in the oviducts), or a male that has produced spermatophores (these are stored in Needham's sac). http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 main nidamental gland anal flap A small flap on the lateral side of the anus. anal valve A small flap on the lateral side of the anus. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 anterior salivary gland Glands on or in the buccal mass that aid in preliminary digestion. Glands on or in the buccal mass that aid in preliminary digestion. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 antitragus Knob that projects inward from the posterior surface of the central depression in the funnel locking-apparatus of some squids. Knob that projects inward from the posterior surface of the central depression in the funnel locking-apparatus of some squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 areolar spot Circular spots, generally light colored but often with complex pigmentation and of unknown function that occur in lines along the aboral surfaces of some arms and portions of the head in some opisthoteuthid cirrates. Circular spots, generally light colored but often with complex pigmentation and of unknown function that occur in lines along the aboral surfaces of some arms and portions of the head in some opisthoteuthid cirrates. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 arm One of the circumoral appendages of cephalopods. In decapodiforms each member of the fourth pair of arms is modified to form a tentacle. circumoral arm One of the circumoral appendages of cephalopods. In decapodiforms each member of the fourth pair of arms is modified to form a tentacle. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 arm formula The relative sizes of arms arranged according to their length. The largest arm is indicated first and the shortest last. An example arm formula is IV>III=II>I. Here arm IV is the largest, followed by arm III which is the same size as arm II and both are larger than arm I. The relative sizes of arms arranged according to their length. The largest arm is indicated first and the shortest last. An example arm formula is IV>III=II>I. Here arm IV is the largest, followed by arm III which is the same size as arm II and both are larger than arm I. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 arm IV In some squids arms IV are much longer and broader than the other arms. ventral arm In some squids arms IV are much longer and broader than the other arms. http://tolweb.org/accessory/Cephalopod_Arms_IV:_Size?acc_id=2017 armature of the arms The grappling structures on the oral surfaces the arms and tentacles, including both suckers and hooks. The grappling structures on the oral surfaces the arms and tentacles, including both suckers and hooks. http://tolweb.org/accessory/Armature_of_Cephalopod_Arms?acc_id=2018 a-seriation Type of cusp arrangement on the rachadian tooth of the radula: multiple cusps seem to move laterally when viewing successive rachadian teeth from adjacent rows. Type of cusp arrangement on the rachadian tooth of the radula: multiple cusps seem to move laterally when viewing successive rachadian teeth from adjacent rows. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 autotomy plane Plane through an arm of an octopod that is predisposed to break when the arm is autotomized. Plane through an arm of an octopod that is predisposed to break when the arm is autotomized. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 basal shelf See cirrate shell terminology. See cirrate shell terminology. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 beak One of the two chitinous jaws of cephalopods. The dorsal beak is the &quot;upper&quot; beak and it inserts within the lower beak to tear tissue with a sissors-like cutting action. http://en.wikipedia.org/wiki/Cephalopod_beak rostrum One of the two chitinous jaws of cephalopods. The dorsal beak is the &quot;upper&quot; beak and it inserts within the lower beak to tear tissue with a sissors-like cutting action. http://tolweb.org/accessory/Cephalopod_Beak_Terminology?acc_id=1980 beak step A ridge on the lower beak where the rostral edge merges onto the lateral wall. In most beaks the merger is smooth. This is a characteristic feature of the Onychoteuthidae. A ridge on the lower beak where the rostral edge merges onto the lateral wall. In most beaks the merger is smooth. This is a characteristic feature of the Onychoteuthidae. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 branchial canal A large opening at the base of each gill lamella and between the primary afferent and efferent blood vessels of the gill. A branchial canal is absent in <strong>Nautilus</strong> , sepioid decapodiformes and cirrate octopods. A large opening at the base of each gill lamella and between the primary afferent and efferent blood vessels of the gill. A branchial canal is absent in <strong>Nautilus</strong> , sepioid decapodiformes and cirrate octopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 brachial crown The combination of arms and tentacles that surround the mouth. The combination of arms and tentacles that surround the mouth. http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 brachial lobe of brain The anterior-most part of the brain located ventral to the esophagus. The large axial nerve cords that run down the centers of the arms connect to this lobe. In octopods this lobe is fused with the rest of the subesophageal mass while in decapodiforms and <strong>Vampyroteuthis</strong> it is separate from the remaining brain. anterior subesophageal mass anterior subesophageal mass of brain The anterior-most part of the brain located ventral to the esophagus. The large axial nerve cords that run down the centers of the arms connect to this lobe. In octopods this lobe is fused with the rest of the subesophageal mass while in decapodiforms and <strong>Vampyroteuthis</strong> it is separate from the remaining brain. http://tolweb.org/accessory/Cephalopod_Brain_Terminology?acc_id=1944 branchial gland Elongate gland adjacent to and paralleling the gill where the gill attaches to the mantle wall. Elongate gland adjacent to and paralleling the gill where the gill attaches to the mantle wall. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 branchial heart A pulsating gland at the base of the gill and through which the afferent blood to the gills flows. It contributes to the blood flow through the gill but also is the site of hemocyanin (the blood respiratory pigment) synthesis. http://en.wikipedia.org/wiki/Branchial_heart Branchial hearts appear to be involved in hemocyanin synthesis http://en.wikipedia.org/wiki/Branchial_heart Branchial hearts appear to be involved in hemocyanin synthesis A pulsating gland at the base of the gill and through which the afferent blood to the gills flows. It contributes to the blood flow through the gill but also is the site of hemocyanin (the blood respiratory pigment) synthesis. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 brachial photophore Photophores located on the arms. photophore at base of gill[http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015]l http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 photophore at base of gill[http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015]l Photophores located on the arms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 brachial pillar A very narrow and elongate anterior region of the head between the eyes and the base of the brachial crown. A branchial pillar is especially well developed in many young cranchiid squid. A very narrow and elongate anterior region of the head between the eyes and the base of the brachial crown. A branchial pillar is especially well developed in many young cranchiid squid. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod brain Medial portion of the central nervous system. It includes the subesophageal and supraesophageal masses but generally does not include the large optic lobes. Medial portion of the central nervous system. It includes the subesophageal and supraesophageal masses but generally does not include the large optic lobes. http://tolweb.org/accessory/Cephalopod_Brain_Terminology?acc_id=1944 brooding Incubation of eggs by the parent. A characteristic feature of incirrate octopods but also found in some squid (e.g., Gonatidae). Incubation of eggs by the parent. A characteristic feature of incirrate octopods but also found in some squid (e.g., Gonatidae). http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 buccal connective Muscular membrane that runs from the buccal support to the base of the adjacent arm. The position of the connective on the fourth arms was recognized in the early 20th century as an important character in interpreting phylogenetic relationships among decapodiforms. Muscular membrane that runs from the buccal support to the base of the adjacent arm. The position of the connective on the fourth arms was recognized in the early 20th century as an important character in interpreting phylogenetic relationships among decapodiforms. http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 buccal crown Umbrella-like structure that surrounds the mouth and is surrounded by the brachial crown. It consists of buccal supports and the buccal membrane. The buccal crown is present in most decapodiforms but absent from all octopodiforms. Umbrella-like structure that surrounds the mouth and is surrounded by the brachial crown. It consists of buccal supports and the buccal membrane. The buccal crown is present in most decapodiforms but absent from all octopodiforms. http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 buccal mass Muscular bulb at the beginning of the digestive system that contains the mouth, beaks, radula and various glands. buccal system Muscular bulb at the beginning of the digestive system that contains the mouth, beaks, radula and various glands. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 buccal membrane The muscular membrane that connects the buccal supports and together with the supports, forms the buccal crown. The pigmentation of the buccal membrane often differs from that of the adjacent oral surfaces of the arms. The muscular membrane that connects the buccal supports and together with the supports, forms the buccal crown. The pigmentation of the buccal membrane often differs from that of the adjacent oral surfaces of the arms. http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 buccal sucker Small but normal suckers that are present on the oral surface of the buccal supports of some decapodiforms (bathyteuthid families, some loliginids and some sepiids). TODO - add buccal membrane support Small but normal suckers that are present on the oral surface of the buccal supports of some decapodiforms (bathyteuthid families, some loliginids and some sepiids). http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 bursa copulatrix A spermatophore receptacle found in some sepiolids and consisting of folded glandular epithelium of the lining of the mantle cavity just anterior to the oviduct opening. A spermatophore receptacle found in some sepiolids and consisting of folded glandular epithelium of the lining of the mantle cavity just anterior to the oviduct opening. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 caecal sac The sac-like, thin-walled posterior portion of the caecum that lacks the internal, ciliated leaflets characteristic of the anterior portion of the caecum. The sac-like, thin-walled posterior portion of the caecum that lacks the internal, ciliated leaflets characteristic of the anterior portion of the caecum. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 caecum Region of the digestive tract of all cephalopods between the stomach and intestine. It is the primary site of food absorption. Region of the digestive tract of all cephalopods between the stomach and intestine. It is the primary site of food absorption. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 calcareous Composed of calcium carbonate (CaCO3) Composed of calcium carbonate (CaCO3) http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 calamus The conical papilla of the end-organ of the hectocotylus in many incirrate octopods. It is located at the distal termination of the sperm groove and usually lies adjacent to the last arm sucker. The conical papilla of the end-organ of the hectocotylus in many incirrate octopods. It is located at the distal termination of the sperm groove and usually lies adjacent to the last arm sucker. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cement body Structure in the spermatophore that allows adhesion of the discharged spermatophore to a female. Structure in the spermatophore that allows adhesion of the discharged spermatophore to a female. http://tolweb.org/accessory/Cephalopod_Spermatophore_Terminology?acc_id=1972 carpal sucker Suckers in the locking apparatus at the base of the club. Suckers in the locking apparatus at the base of the club. http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 carpal knob Knobs in the locking apparatus at the base of the club. Knobs in the locking apparatus at the base of the club. http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 carpus The region of the tentacle that contains the carpal locking-apparatus The term 'carpus' is most appropriate where the carpal locking-apparatus is separate from the manus (e.g., Onychoteuthis spp.). The region of the tentacle that contains the carpal locking-apparatus http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 The term 'carpus' is most appropriate where the carpal locking-apparatus is separate from the manus (e.g., Onychoteuthis spp.). carpal locking apparatus The region at the base of the tentacular club which has small suckers and knobs. The carpal suckers and knobs of one tentacle can adhere to their counterparts on the opposite tentacle which, thereby, holds the clubs together at their bases. proximal locking-apparatus The region at the base of the tentacular club which has small suckers and knobs. The carpal suckers and knobs of one tentacle can adhere to their counterparts on the opposite tentacle which, thereby, holds the clubs together at their bases. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cartilaginous tubercle Rigid cartilagenous spines, with one or several cusps, in the skin of certain squids and octopods. Rigid cartilagenous spines, with one or several cusps, in the skin of certain squids and octopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalic cartilage Cartilage surrounding the posterior part of the brain of cephalopods and generally encompasing the statocysts. Anteriorly the cartilage thins into facia making a well-defined limit difficult to establish. The cartilage has a large central foramen through which the esophagus passes and minor foramina for nerves and blood vessels. Cartilage surrounding the posterior part of the brain of cephalopods and generally encompasing the statocysts. Anteriorly the cartilage thins into facia making a well-defined limit difficult to establish. The cartilage has a large central foramen through which the esophagus passes and minor foramina for nerves and blood vessels. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalic vein Large vein that drains the head and lies on the ventral surface of the visceral sac, along side or dorsal to the intestine. The cephalic vein terminates by dividing into the two vena cavae, each of which passes through the &quot;kidney&quot; (nephridium), the branchial heart and into the gill. Large vein that drains the head and lies on the ventral surface of the visceral sac, along side or dorsal to the intestine. The cephalic vein terminates by dividing into the two vena cavae, each of which passes through the &quot;kidney&quot; (nephridium), the branchial heart and into the gill. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 chitin A mucopolysaccharide that is the major component of the cuticle (exoskeleton) of arthropods and a major component of the gladius and beaks of cephalopods (also found in many other phyla). TODO - replace with uberon A mucopolysaccharide that is the major component of the cuticle (exoskeleton) of arthropods and a major component of the gladius and beaks of cephalopods (also found in many other phyla). http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 chromatophore A pigment cell. In cephalopods the chromatophore is part of an organ composed of numerous cells including many slender muscle cells. Under nervous control, the muscles rapidly expand and flatten the pigment cell which spreads the pigment over a relatively broad area. A pigment cell. In cephalopods the chromatophore is part of an organ composed of numerous cells including many slender muscle cells. Under nervous control, the muscles rapidly expand and flatten the pigment cell which spreads the pigment over a relatively broad area. http://tolweb.org/accessory/Cephalopod_Chromatophore?acc_id=2038 circularis muscle A circular muscle found in the suckers of most sepioids and loliginids that surrounds the sucker at the base of the horny ring. A circular muscle found in the suckers of most sepioids and loliginids that surrounds the sucker at the base of the horny ring. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cirrate male reproductive system The male reproductive tract of cirrates does not make typical spermatophores. As a result, the tract differs greatly from that of other coleoid cephalopods. The male reproductive tract of cirrates does not make typical spermatophores. As a result, the tract differs greatly from that of other coleoid cephalopods. http://tolweb.org/accessory/Cirrate_Male_Reproductive_Tract?acc_id=1486 cirrate octopod measurements Measurements used in describing cirrates differ somewhat from other cephalopods. Measurements used in describing cirrates differ somewhat from other cephalopods. http://tolweb.org/accessory/Cirrate_Measurements_and_Indices?acc_id=1487 arm cirrus Muscular, conical structures that arise from the sides of the arms of some octopodiforms. Cirri are homologous with trabeculae of decapodiforms. Muscular, conical structures that arise from the sides of the arms of some octopodiforms. Cirri are homologous with trabeculae of decapodiforms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod coelom An internal body cavity of mesodermal orgin that is lined by an epithelium. Cephalopods have two coeloms, the visceropericardial coelom and the nephridial coelom. An internal body cavity of mesodermal orgin that is lined by an epithelium. Cephalopods have two coeloms, the visceropericardial coelom and the nephridial coelom. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 collar Muscular structure extending from the nuchal cartilage to the funnel that forms a one-way valve that lets water enter the mantle cavity but closes as the mantle contracts thereby forcing exhalent water out through the funnel. Muscular structure extending from the nuchal cartilage to the funnel that forms a one-way valve that lets water enter the mantle cavity but closes as the mantle contracts thereby forcing exhalent water out through the funnel. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 photophore color filter Structures that alter the color of bioluminescent light. Structures that alter the color of bioluminescent light. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 conus The cup or cone-shaped posterior end of the gladius or cuttlebone of some cephalopods. It is not formed by the infolding of the vanes. conus flag cone The cup or cone-shaped posterior end of the gladius or cuttlebone of some cephalopods. It is not formed by the infolding of the vanes. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 conus field Lateral portions of the gladius that are continuous posteriorly with the primary conus or its remnant and laterally with the edges of the vanes. Lateral portions of the gladius that are continuous posteriorly with the primary conus or its remnant and laterally with the edges of the vanes. http://tolweb.org/accessory/Cephalopod_Gladius_Terminology?acc_id=1955 cornea Smooth, thin, turgid, transparent skin without muscles that covers the eyes to protect the eye lenses of incirrate octopods and some decapodiforms. todo - reconcile with metazoan def in uberon Smooth, thin, turgid, transparent skin without muscles that covers the eyes to protect the eye lenses of incirrate octopods and some decapodiforms. http://tolweb.org/accessory/Cephalopod_Cornea?acc_id=2057 counterillumination The production of bioluminescent light by an animal to conceal its silhouette against the downwelling light. The process can allow an aninmal to become virtually invisible under dim directional light. The production of bioluminescent light by an animal to conceal its silhouette against the downwelling light. The process can allow an aninmal to become virtually invisible under dim directional light. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 crest of hectocotylus In loliginids a protective membrane on the hectocotylized arm can fuse with the adjacent suckerless papillae (former sucker stalks) to form a crest on the dorsal or ventral margins such that the original form of the papillae is obscured. In loliginids a protective membrane on the hectocotylized arm can fuse with the adjacent suckerless papillae (former sucker stalks) to form a crest on the dorsal or ventral margins such that the original form of the papillae is obscured. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 crop Expansion (i.e. a broading or a side pocket) of the esophagus for storing food. Present in <strong>Nautilus</strong> and most octopodiforms. Expansion (i.e. a broading or a side pocket) of the esophagus for storing food. Present in <strong>Nautilus</strong> and most octopodiforms. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 ctenoglossan radula Radula in which the lateral teeth are multicuspid. Radula in which the lateral teeth are multicuspid. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cupped coil The peculiar posterior end of the histioteuthis gladius that is rolled into a coil. The peculiar posterior end of the histioteuthis gladius that is rolled into a coil. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cusp A point or projection on a tooth of the radula or on a cartilagenous tubercule in the skin. todo - add subclasses for radula or skin A point or projection on a tooth of the radula or on a cartilagenous tubercule in the skin. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cuttlebone Predominately calcareous internal shell of sepiids (cuttlefish). It is composed of many chambers and functions primarily as a buoyancy organ. The cuttlebone is homologous with the phragmocone of <strong>Spirula</strong>, <strong>Nautilus</strong> and many fossil cephalopods. sepion A calcareous oblong supporting part of the mantle of cuttlefishes http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A calcareous oblong supporting part of the mantle of cuttlefishes Predominately calcareous internal shell of sepiids (cuttlefish). It is composed of many chambers and functions primarily as a buoyancy organ. The cuttlebone is homologous with the phragmocone of <strong>Spirula</strong>, <strong>Nautilus</strong> and many fossil cephalopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 dactylus The narrow, distal portion of the tentacular club of most decapodiforms. In squids the dactylus is usually characterized by the ventral expansion of the club and the absence of a dorsal protective membrane. The narrow, distal portion of the tentacular club of most decapodiforms. In squids the dactylus is usually characterized by the ventral expansion of the club and the absence of a dorsal protective membrane. http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 demersal Living close to the ocean floor. Living close to the ocean floor. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 demibranch A half-gill formed from all the primary gill lamellae on one side of the gill. A half-gill formed from all the primary gill lamellae on one side of the gill. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 diel vertical migration Vertical animal migration during twilight. Many mesopelagic animals migrate at sunrise from near-surface waters to spend the day at greater depths then return to shallow depths at sunset where they spend the night. Some animals migrate vertically over 1000 m others migrate less than 100 m. Vertical animal migration during twilight. Many mesopelagic animals migrate at sunrise from near-surface waters to spend the day at greater depths then return to shallow depths at sunset where they spend the night. Some animals migrate vertically over 1000 m others migrate less than 100 m. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 digestive gland Primary organ in cephalopods that secretes digestive enzymes. It is also important in absorption and excretion. Primary organ in cephalopods that secretes digestive enzymes. It is also important in absorption and excretion. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 digestive gland duct appendages (digda) Outpockets of the ducts leading from the digestive gland that are covered with glandular epithelium. Outpockets of the ducts leading from the digestive gland that are covered with glandular epithelium. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 distal oviduct An extension of the oviducal gland in octopods that often has the same general appearance as the oviduct proximal to the oviducal gland. An extension of the oviducal gland in octopods that often has the same general appearance as the oviduct proximal to the oviducal gland. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 doratopsis stage Peculiar paralarval stage of the Chiroteuthidae. The paralarva is very slender and can reach a large size. doratopsis Peculiar paralarval stage of the Chiroteuthidae. The paralarva is very slender and can reach a large size. http://tolweb.org/accessory/Chiroteuthid_Doratopsis_Stage?acc_id=656 dorsal mantle cavity Dorsal portion of the mantle cavity that completely surrounds the viscera in octopods. In decapodiforms and <strong>Vampyroteuthis</strong> the viscera are fused to the mantle and shell dorsally and the mantle cavity lies lateral and ventral to the viscera. Dorsal portion of the mantle cavity that completely surrounds the viscera in octopods. In decapodiforms and <strong>Vampyroteuthis</strong> the viscera are fused to the mantle and shell dorsally and the mantle cavity lies lateral and ventral to the viscera. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 efferent vein vein carrying blood away from an organ. todo - mireot http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 vein carrying blood away from an organ. efferent nerve nerve carring impulses away from the brain or specific ganglia. TODO - mireot uberon http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nerve carring impulses away from the brain or specific ganglia. egg mass A large number of eggs encapsulated in a gelatinous matrix or a large number of such structures that are attached together. The egg mass of an oceanic squid can be a large, fragile gelatinous ball carrying many thousands of eggs. In contrast, the egg mass of a neritic squid (loliginid) can be composed of very tough capsules each carrying several hundred eggs with many capsules attached together at their bases and to the substrate to form an egg mass. egg case todo - mireot A large number of eggs encapsulated in a gelatinous matrix or a large number of such structures that are attached together. The egg mass of an oceanic squid can be a large, fragile gelatinous ball carrying many thousands of eggs. In contrast, the egg mass of a neritic squid (loliginid) can be composed of very tough capsules each carrying several hundred eggs with many capsules attached together at their bases and to the substrate to form an egg mass. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 ejaculatory apparatus Portion of the spermatophore involved in the eversion of the sperm mass. Portion of the spermatophore involved in the eversion of the sperm mass. http://tolweb.org/accessory/Cephalopod_Spermatophore_Terminology?acc_id=1972 end organ The tip of the hectocotylus in incirrate octopods consisting of a calamus and ligula presumably for manipulating the spermatophore during copulation. end organ of hectocotylus terminal organ The tip of the hectocotylus in incirrate octopods consisting of a calamus and ligula presumably for manipulating the spermatophore during copulation. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 epithelial pigmentation The pigmentation that is contained in epithelial cells which are unable to change their shape. Color in most cephalopods, however, is due to pigment granules that are contained in specialized organs, the chromatophore organs, that can change shape rapidly under nervous control. See chromatophores. The pigmentation that is contained in epithelial cells which are unable to change their shape. Color in most cephalopods, however, is due to pigment granules that are contained in specialized organs, the chromatophore organs, that can change shape rapidly under nervous control. See chromatophores. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 eye pore Small pore just anterior to the cornea in some decapodiforms. The pore is the remnant of the large eye opening of oegopsids and connects fluid between the lens and the cornea with the exterior environment. Small pore just anterior to the cornea in some decapodiforms. The pore is the remnant of the large eye opening of oegopsids and connects fluid between the lens and the cornea with the exterior environment. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 eyelid sinus Indentation, often complex, of the anterior margin of the eyelid. optic sinus orbital sinus Indentation, often complex, of the anterior margin of the eyelid. http://tolweb.org/accessory/Cephalopod_Eyelid_Sinus?acc_id=1994 fin angle The angle between the longitudinal axis of the mantle and the posterior border of one fin. The angle between the longitudinal axis of the mantle and the posterior border of one fin. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 fin attachment A fin attaches to the shell, the mantle, the opposite fin or some combination of these. A fin attaches to the shell, the mantle, the opposite fin or some combination of these. http://tolweb.org/accessory/Cephalopod_Fin_Attachment?acc_id=1974 fin cartilage Cartilage associated with the fins of all fin-bearing cephalopods. Cartilage associated with the fins of all fin-bearing cephalopods. http://tolweb.org/accessory/Cephalopoda_Fin_Cartilage?acc_id=1969 fin length This measurement varies in different major taxa. This measurement varies in different major taxa. http://tolweb.org/accessory/Cephalopod_Fin_Length?acc_id=2055 fin lobe The portion of a fin that extends anteriorly from the fin's anterior point of attachment or posteriorly from the fin's posterior point of attachment. This is often called the &quot;free&quot; lobe. The portion of a fin that extends anteriorly from the fin's anterior point of attachment or posteriorly from the fin's posterior point of attachment. This is often called the &quot;free&quot; lobe. http://tolweb.org/accessory/Composite_Squid?acc_id=1936 fin position Fins are located anterior to the termination of the muscular mantle (subterminal position) or mostly posterior to it (terminal position) or somewhere inbetween. Fins are located anterior to the termination of the muscular mantle (subterminal position) or mostly posterior to it (terminal position) or somewhere inbetween. http://tolweb.org/accessory/Cephalopod_Fin_Position?acc_id=2041 fin shape Fins are classified, somewhat arbitrarily, by their shape as Sagittate, Rhomboid, Circular/Elliptical, Lanceolate, Ear-shaped, Ribbed, Lobate or skirt-like. Fins are classified, somewhat arbitrarily, by their shape as Sagittate, Rhomboid, Circular/Elliptical, Lanceolate, Ear-shaped, Ribbed, Lobate or skirt-like. http://tolweb.org/accessory/Decapodiform_Fin_Shapes?acc_id=2029 fin The pair of muscular &quot;wings&quot; used in swimming and located along the dorsolateral surface of the mantle, usually in its posterior half . A muscular flap that arises along the lateral or dorsolateral surface of the mantle of some cephalopods. Used for locomotion, steering and stabilization The pair of muscular &quot;wings&quot; used in swimming and located along the dorsolateral surface of the mantle, usually in its posterior half . http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 foveola Transverse membranous fold of skin that forms a pocket in the funnel groove of ommastrephid squids. Transverse membranous fold of skin that forms a pocket in the funnel groove of ommastrephid squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 obsolete free fin lobe The portion of a fin that extends anteriorly from the fin's anterior point of attachment or posteriorly from the fin's posterior point of attachment. true The portion of a fin that extends anteriorly from the fin's anterior point of attachment or posteriorly from the fin's posterior point of attachment. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 funnel The ventral, muscular tube through which water is expelled from the mantle cavity. The funnel is thought to have evolved from the molluscan foot. http://en.wikipedia.org/wiki/Siphon_%28mollusc%29 siphon hyponome The ventral, muscular tube through which water is expelled from the mantle cavity. The funnel is thought to have evolved from the molluscan foot. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 funnel groove Depression on the posteroventral surface of the head in which the funnel lies. Most distinct in some squids. Depression on the posteroventral surface of the head in which the funnel lies. Most distinct in some squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 funnel-mantle locking-apparatus The funnel component (generally cartilagenous) of a locking mechanism between the funnel and mantle. This lock, depending on its form, can either limit all movement between the funnel and mantle or allow just anterior-posterior slipping. See Mantle locking-apparatus. funnel locking apparatus http://en.wikipedia.org/wiki/Funnel%E2%80%93mantle_locking_apparatus funnel locking-cartilage The funnel component (generally cartilagenous) of a locking mechanism between the funnel and mantle. This lock, depending on its form, can either limit all movement between the funnel and mantle or allow just anterior-posterior slipping. See Mantle locking-apparatus. http://tolweb.org/accessory/Cephalopod_Funnel_Locking-Apparatus?acc_id=1973 funnel organ The glandular structures on the internal dorsal and ventral surfaces of the funnel. The glandular structures on the internal dorsal and ventral surfaces of the funnel. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 funnel retractor muscle Large muscles that attach to the corners of the funnel and run posteriorly to attach to the sides of the shell sac (generally near the base of the gills) or, in a some species, insert on the mantle wall. Large muscles that attach to the corners of the funnel and run posteriorly to attach to the sides of the shell sac (generally near the base of the gills) or, in a some species, insert on the mantle wall. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 funnel valve Semicircular muscular flap extending from and continuous with the posterodorsal funnel wall. The funnel valve apparently acts as a one-way valve to prevent inspiration of water through the funnel when mantle expansion draws in water to pass over the gills for respiration. A funnel valve in absent in octopods and some squids. Semicircular muscular flap extending from and continuous with the posterodorsal funnel wall. The funnel valve apparently acts as a one-way valve to prevent inspiration of water through the funnel when mantle expansion draws in water to pass over the gills for respiration. A funnel valve in absent in octopods and some squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 gill Primary organ for the exchange of respiratory gases between blood and sea water. Primary organ for the exchange of respiratory gases between blood and sea water. http://tolweb.org/articles/?article_id=4200 gill lamella A complex gill leaflet that extends perpendicular to the axis of the gill and contains the respiratory epithelium. The lamella contains further subdivisions, the secondary and tertiary lamellae. In octopods the number of lamellae is a taxonomic character. gill lamellae primary lamella A complex gill leaflet that extends perpendicular to the axis of the gill and contains the respiratory epithelium. The lamella contains further subdivisions, the secondary and tertiary lamellae. In octopods the number of lamellae is a taxonomic character. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 gladius The chitinous remnant of the shell in many decapodiforms and <strong>Vampyroteuthis</strong>. It is generally feather-shaped and lies in the dorsal mid-line of the body along the full length of the mantle. The gladius lies within the shell sac which secretes it. The gladius, found in many decapodiforms and Vampyroteuthis, is an internal remnant of the originally external shell of cephalopods. Other cephalopods have other types of shell remnants. The gladius lies within a sac, the shell sac, which secretes it and to which various muscles attach. The gladius has a variety of shapes and thicknesses, is composed mostly of chitin and is located in the dorsal midline and generally extends the full length of the mantle. Conus field - The lateral portion of the gladius that is continuous posteriorly with the primary conus or its remnant and laterally with the edges of the vanes. The conus field is differentiated from the vanes by longitudinal lines and marked changes in the orientation of the growth lines. A conus field is generally lacking if the gladius lacks a primary conus. http://tolweb.org/tree/ToLimages/Gladius.jpg gladii internal gladius pen http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf The chitinous remnant of the shell in many decapodiforms and <strong>Vampyroteuthis</strong>. It is generally feather-shaped and lies in the dorsal mid-line of the body along the full length of the mantle. The gladius lies within the shell sac which secretes it. http://tolweb.org/accessory/Cephalopod_Gladius_Terminology?acc_id=1955 half-orange gill Gills of some cirrate octopods with broad primary lamellae whose arrangement looks similar to half of an orange. Gills of some cirrate octopods with broad primary lamellae whose arrangement looks similar to half of an orange. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 hatchling stage Young cephalopod that has just hatched from the egg. hatchling todo - stages Young cephalopod that has just hatched from the egg. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 head-mantle fusion . . http://tolweb.org/accessory/Cephalopod_Dorsal_Head-Mantle_Fusion?acc_id=2020 hectocotylus Taxon notes: Among Decapodiformes (ten-limbed cephalopods), generally either one or both of arms IV are hectocotylized; in incirrate octopuses it is one of arm pair III.[1] Many coleoids lack hectocotyli altogether.[1] The shape of the tip of the hectocotylus has been much used in octopod systematics. In many species it is considerably elaborated. However, in the males of some species, such as the Seven-arm Octopus (Haliphron atlanticus), the hectocotylus develops in an inconspicuous sac in front of the right eye that gives the male the appearance of having only seven arms The modified arm in males used for transferring spermatophores to the female. In incirrates the hectocotylus is one member of arm pair III while in decapodiforms frequently one or both members of arm pair IV are modified. Many species of coleoid cephalopods lack a hectocotylus. http://upload.wikimedia.org/wikipedia/commons/thumb/9/99/Hectocotylized_arm_of_a_squid.jpg/189px-Hectocotylized_arm_of_a_squid.jpg hectoctylized arm Wikipedia The modified arm in males used for transferring spermatophores to the female. In incirrates the hectocotylus is one member of arm pair III while in decapodiforms frequently one or both members of arm pair IV are modified. Many species of coleoid cephalopods lack a hectocotylus. Wikipedia:Hectocotylus http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 heterodont radula Radula with more than one cusp on the rhachidian tooth and, usually, the first lateral tooth. Radula with more than one cusp on the rhachidian tooth and, usually, the first lateral tooth. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 homodont radula Radula with a single cusp on all teeth. Radula with a single cusp on all teeth. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 obsolete holotype The single specimen designated to represent a species name. It is an international standard of reference that provides objectivity for the species name. consider obsoleting true The single specimen designated to represent a species name. It is an international standard of reference that provides objectivity for the species name. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 hook Horny structure that looks like a single claw and is derived from the inner sucker ring in some decapodiforms. Hooks may be found on the arms and/or tentacular clubs of squid. The relative advantage of hooks vs toothed rings is uncertain. ontogenically derived from suckers on arms and/or clubs of some oegopsids Horny structure that looks like a single claw and is derived from the inner sucker ring in some decapodiforms. Hooks may be found on the arms and/or tentacular clubs of squid. The relative advantage of hooks vs toothed rings is uncertain. http://tolweb.org/accessory/Cephalopod_Hooks?acc_id=1983 horizontal arm septum Septa, extending the length of the arm (i.e., parallel to the arm axis), that roughly divides the arm into oral and aboral regions. This feature is a characteristic of the arms of cirrate octopods and the incirrate octopods of the family Bolitaenidae. The functional significance is unknown. todo - mireot Septa, extending the length of the arm (i.e., parallel to the arm axis), that roughly divides the arm into oral and aboral regions. This feature is a characteristic of the arms of cirrate octopods and the incirrate octopods of the family Bolitaenidae. The functional significance is unknown. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sucker ring Suckers of decapodiforms have two types of hard, horny rings. One, the inner ring, lies around the inner walls of the acetabulum and often bears teeth. The other, the outer ring, is composed of numerous platelets and lies on the surface of the infundibulum. horny ring of sucker A chitinous ring that encircles a sucker. Found in cuttlefishes and squids Suckers of decapodiforms have two types of hard, horny rings. One, the inner ring, lies around the inner walls of the acetabulum and often bears teeth. The other, the outer ring, is composed of numerous platelets and lies on the surface of the infundibulum. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 inferior frontal lobe system A system of lobes in octopod brains: the paired posterior buccal, lateral inferior frontal and subfrontal lobes, and the single median inferior frontal lobe. They form a functional unit concerned with the chemotactile information from the arms and its use. A system of lobes in octopod brains: the paired posterior buccal, lateral inferior frontal and subfrontal lobes, and the single median inferior frontal lobe. They form a functional unit concerned with the chemotactile information from the arms and its use. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 infundibilum The siphon of Cephalopoda Exposed flat outer ring that forms the adhesive surface of the sucker The siphon of Cephalopoda http://thinkexist.com/dictionary/meaning/infundibulum/ http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Exposed flat outer ring that forms the adhesive surface of the sucker ink sac Organ composed of a gland that secretes ink, a sac that stores ink and a duct that connects it to the rectum. The ink sac generally appears black from the outside although it may be covered by silvery tissue in some species. Organ composed of a gland that secretes ink, a sac that stores ink and a duct that connects it to the rectum. The ink sac generally appears black from the outside although it may be covered by silvery tissue in some species. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 jet propulsion This is the typical method of fast swimming in cephalopods. During jet propulsion, water is taken into the mantle cavity by expansion of the mantle. Water enters laterally at the sides of the head, passes over the collar, over the gills and leaves through the funnel when the mantle contracts. Mantle contraction causes water pressure in the mantle cavity to increase and forces the collar against the mantle wall. The collar, therefore, acts as a one-way valve to prevents water from leaving by the same route that it entered. The ventral wall of the funnel presses against the mantle during mantle contraction and seals that opening as well. Water, therefore, is forced through the funnel and out the funnel orifice. The force provided by the water leaving the funnel can be directed by movements of the funnel. Typically the water leaves the funnel anteriorly causing the cephalopod to jet backward (e.g. when escaping a predator) or the funnel is directed posteriorly causing the water to leave posteriorly and the animal to jet anteriorly (e.g. when pouncing on a prey). Some species are able to glide via this method. This is the typical method of fast swimming in cephalopods. During jet propulsion, water is taken into the mantle cavity by expansion of the mantle. Water enters laterally at the sides of the head, passes over the collar, over the gills and leaves through the funnel when the mantle contracts. Mantle contraction causes water pressure in the mantle cavity to increase and forces the collar against the mantle wall. The collar, therefore, acts as a one-way valve to prevents water from leaving by the same route that it entered. The ventral wall of the funnel presses against the mantle during mantle contraction and seals that opening as well. Water, therefore, is forced through the funnel and out the funnel orifice. The force provided by the water leaving the funnel can be directed by movements of the funnel. Typically the water leaves the funnel anteriorly causing the cephalopod to jet backward (e.g. when escaping a predator) or the funnel is directed posteriorly causing the water to leave posteriorly and the animal to jet anteriorly (e.g. when pouncing on a prey). Some species are able to glide via this method. Wikipedia:Ommastrephidae http://tolweb.org/accessory/Cephalopod_Jet_Propulsion?acc_id=2060 keel A muscular vane that extends dorsally from the aboral surface of the tentacular club of many species. club swimming membrane A muscular vane that extends dorsally from the aboral surface of the tentacular club of many species. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 lateral funnel adductor muscle Muscles that support the lateral attachment of the funnel to the head. Muscles that support the lateral attachment of the funnel to the head. http://tolweb.org/accessory/Cephalopod_Lateral_Funnel_Adductor?acc_id=1963 lateral membrane of arm IV The keels of arms IV are offset laterally and often enlarged to fully or partially conceal, protect or encase the adjacent tentacles. The latter function is most fully developed in the chiroteuthids and mastigoteuthids. tentacular sheath The keels of arms IV are offset laterally and often enlarged to fully or partially conceal, protect or encase the adjacent tentacles. The latter function is most fully developed in the chiroteuthids and mastigoteuthids. http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 http://tolweb.org/accessory/Cephalopod_Brachial_and_Buccal_Crowns?acc_id=1957 tentacular sheath lateral-line analogue Sensory structure analogous to the lateral-line of fishes. The lateral-line analogue, which senses vibrations transmitted by the seawater, is located along a series of lines on the dorsal surface of the head with some sensory cells extending onto the bases of the arms. Sensory structure analogous to the lateral-line of fishes. The lateral-line analogue, which senses vibrations transmitted by the seawater, is located along a series of lines on the dorsal surface of the head with some sensory cells extending onto the bases of the arms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 photophore lens Structure in a photophore that can focus or disperse bioluminescent light. Structure in a photophore that can focus or disperse bioluminescent light. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 light guides Structures in a photophores that direct light via internal reflection. Structures in a photophores that direct light via internal reflection. http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 ligula The portion of the sucker-free end-organ of the hectocotylus of many incirrate octopods that is generally elongate and longitudinally grooved. The ligula in octopodids is often spatulate in shape with transverse ridges. Spermatophores transferred along the sperm groove of the hectocotylus presumably enter the ligula which assists in their transfer to the female. Details of the use of the ligula are unknown. The portion of the sucker-free end-organ of the hectocotylus of many incirrate octopods that is generally elongate and longitudinally grooved. The ligula in octopodids is often spatulate in shape with transverse ridges. Spermatophores transferred along the sperm groove of the hectocotylus presumably enter the ligula which assists in their transfer to the female. Details of the use of the ligula are unknown. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 lip Two concentric muscular, glandular rings of skin that surround the mouth and beaks. Two concentric muscular, glandular rings of skin that surround the mouth and beaks. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 living chamber Largest and most recently formed chamber of the <strong>Nautilus</strong> shell (also refers to numerous shells of fossil cephalopods) in which the animal resides. Largest and most recently formed chamber of the <strong>Nautilus</strong> shell (also refers to numerous shells of fossil cephalopods) in which the animal resides. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 locking-apparatus of tentacular stalk Continuation of the carpal locking apparatus down the oral face of the tentacular stalk. Continuation of the carpal locking apparatus down the oral face of the tentacular stalk. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 mantle locking-apparatus The mantle component (usually fibrous) of the structure that &quot;locks&quot; the mantle to the funnel. This lock, depending on its form, can either restrict all movement between the funnel and mantle or allow just an anterior/posterior slippage. mantle locking cartilage The mantle component (usually fibrous) of the structure that &quot;locks&quot; the mantle to the funnel. This lock, depending on its form, can either restrict all movement between the funnel and mantle or allow just an anterior/posterior slippage. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 mantle length (ML) - The standard measure of length in coleoid cephalopods. In decapodiforms ML is measured along the dorsal midline from the mantle margin to the posterior tip of the body, excluding long tails. In octopodiforms ML is measured from a line joining the midpoint of the eyes rather than the anterior mantle margin since the latter is obscured by the head/mantle fusion. (ML) - The standard measure of length in coleoid cephalopods. In decapodiforms ML is measured along the dorsal midline from the mantle margin to the posterior tip of the body, excluding long tails. In octopodiforms ML is measured from a line joining the midpoint of the eyes rather than the anterior mantle margin since the latter is obscured by the head/mantle fusion. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 mantle septum A anteroposterior septum that connects the mantle wall with the visceral sac along the midline in the ventral mantle cavity. The ventral mantle artery runs along the anterior end of this septum. abdominal septum A anteroposterior septum that connects the mantle wall with the visceral sac along the midline in the ventral mantle cavity. The ventral mantle artery runs along the anterior end of this septum. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 manus of tentacular club The proximal, broad portion of the tentacular club. In squids the manus is generally equally expanded on both lateral margins and often includes enlarged suckers. manus The proximal, broad portion of the tentacular club. In squids the manus is generally equally expanded on both lateral margins and often includes enlarged suckers. http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 mature stage In cephalopods this term refers to sexual maturity which is determined for females by the presence of ova (mature eggs) free in the coelom or oviducts, and for males by the presence of spermatophores in Needham's sac. In cephalopods this term refers to sexual maturity which is determined for females by the presence of ova (mature eggs) free in the coelom or oviducts, and for males by the presence of spermatophores in Needham's sac. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 molluscan foot A major structure in molluscan morphology. In gastropods the foot is the muscular sole that the animal crawls with. In cephalopods the funnel, and possibly the arms and tentacles are derived from the molluscan foot. The evolutionary origin of the latter is still uncertain. They may represent outgrowths of the head (favored by anatomical evidence: nerve connections) or modifications of the molluscan foot that have migrated around the mouth (favored by embryological evidence: migration of arm primordia). foot A major structure in molluscan morphology. In gastropods the foot is the muscular sole that the animal crawls with. In cephalopods the funnel, and possibly the arms and tentacles are derived from the molluscan foot. The evolutionary origin of the latter is still uncertain. They may represent outgrowths of the head (favored by anatomical evidence: nerve connections) or modifications of the molluscan foot that have migrated around the mouth (favored by embryological evidence: migration of arm primordia). http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 neck The region separating the posterior end of the cephalic cartilage and the collar. Only those cephalopods with elongate heads (e.g., chiroteuthids) have distinct necks. The region separating the posterior end of the cephalic cartilage and the collar. Only those cephalopods with elongate heads (e.g., chiroteuthids) have distinct necks. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 Needham's sac In males, the large storage sac for spermatophores that is an expanded region of the genital duct at the base of the penis. spermatophoric sac spermatophore sac In males, the large storage sac for spermatophores that is an expanded region of the genital duct at the base of the penis. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nephridial coelom The cavity of the renal sac. It connects with the exterior via the renal pore and with the visceropericardial coelom via a pair of slender ducts from the latter. The cavity of the renal sac. It connects with the exterior via the renal pore and with the visceropericardial coelom via a pair of slender ducts from the latter. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nidamental gland Large glandular structures found in decapodiforms and Nautilus that lie in and open directly into the mantle cavity. The glands are composed of numerous lamellae that are involved in secretion of egg cases or the jelly of egg masses. Large glandular structures found in decapodiforms and Nautilus that lie in and open directly into the mantle cavity. The glands are composed of numerous lamellae that are involved in secretion of egg cases or the jelly of egg masses. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nuchal cartilage See nuchal locking-apparatus. An oblong cartilaginou plate on the posterodorsal surface of the head of most squids and cuttlefishes that articulates with a complementary structure on the inner surface of the anterodorsal surface of the mantle See nuchal locking-apparatus. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf An oblong cartilaginou plate on the posterodorsal surface of the head of most squids and cuttlefishes that articulates with a complementary structure on the inner surface of the anterodorsal surface of the mantle nuchal crest See Occipital crest. See Occipital crest. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nuchal membrane See occipital membrane. See occipital membrane. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nuchal organ Small sensory organ with photoreceptor-like sensory cells that is located in the nuchal region of, apparently, all coleoid cephalopods. Small sensory organ with photoreceptor-like sensory cells that is located in the nuchal region of, apparently, all coleoid cephalopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 nuchal region The posterodorsal region of the head and the area immediately posterior to it. Normally covered by anterior mantle wall The posterodorsal region of the head and the area immediately posterior to it. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 occipital crest Prominent transverse ridge in most decapodiforms that extends across the dorsal and along the lateral surfaces of the head at its posterior end. Prominent transverse ridge in most decapodiforms that extends across the dorsal and along the lateral surfaces of the head at its posterior end. http://tolweb.org/accessory/Cephalopod_Occipital_Region_Terminology?acc_id=1970 occipital fold Fixed folds of the head integument that adjoin the nuchal crest posteriorly and are perpendicular to it. The function of the folds is uncertain. Fixed folds of the head integument that adjoin the nuchal crest posteriorly and are perpendicular to it. The function of the folds is uncertain. http://tolweb.org/accessory/Cephalopod_Occipital_Region_Terminology?acc_id=1970 occipital membrane A membrane connecting the main occipital folds at their posterior ends. TODO - mireot membrane A membrane connecting the main occipital folds at their posterior ends. http://tolweb.org/accessory/Cephalopod_Occipital_Region_Terminology?acc_id=1970 cephalopod olfactory organ A chemosensory organ present on the head or neck in all coleoid cephalopods. The olfactory organ is thought to be the homologue of the rhinophore of <strong>Nautilus</strong>. rhinophore A chemosensory organ present on the head or neck in all coleoid cephalopods. The olfactory organ is thought to be the homologue of the rhinophore of <strong>Nautilus</strong>. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 ontogenetic descent The progressively descent into a deeper-water habitat as a cephalopod grows older and larger. This distribution pattern is a particularly common in many pelagic cranchiid squids. The progressively descent into a deeper-water habitat as a cephalopod grows older and larger. This distribution pattern is a particularly common in many pelagic cranchiid squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 optic lobe of brain Large lobes of the brain associated with the eyes. In octopods and some squids the optic lobes may be separated from the rest of the brain by an optic stalk of varying length. In <strong>Octopus</strong> the optic lobes contain 92 million cells compared with only 42 million in the main central mass of the brain (J. Young, 1963). editor note: check if this is part of the brain optic lobe Large lobes of the brain associated with the eyes. In octopods and some squids the optic lobes may be separated from the rest of the brain by an optic stalk of varying length. In <strong>Octopus</strong> the optic lobes contain 92 million cells compared with only 42 million in the main central mass of the brain (J. Young, 1963). http://tolweb.org/accessory/Cephalopod_Brain_Terminology?acc_id=1944 outer statocyst capsule Capsule in octopodiforms that is embedded in the cephalic cartilage and contains the statocyst capsule. Capsule in octopodiforms that is embedded in the cephalic cartilage and contains the statocyst capsule. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 oviducal gland Gland surrounding the end of the primary oviduct and responsible for secreting some of the external coatings over spawned eggs or, in incirrates, the cement at the base of the chorion stalk . In octopods it also acts as a spermatheca, and is located around the apparent middle of the oviduct. However, the &quot;distal oviduct&quot; is actually an extension of the oviducal gland. Gland surrounding the end of the primary oviduct and responsible for secreting some of the external coatings over spawned eggs or, in incirrates, the cement at the base of the chorion stalk . In octopods it also acts as a spermatheca, and is located around the apparent middle of the oviduct. However, the &quot;distal oviduct&quot; is actually an extension of the oviducal gland. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 paralarval stage First free-living, cephalopod life-history stage that is found in the near-surface plankton and that differs in both morphology and vertical distribution from older juveniles. First free-living, cephalopod life-history stage that is found in the near-surface plankton and that differs in both morphology and vertical distribution from older juveniles. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod penis The long, muscular end of the male gonoduct that assists in transfering spermatophores to the female. Apparently, in species with a hectocotylus, the penis transfers spermatophores to the hectocotylus which in turn transfers them to the female. In species without a hectocotylus, the penis is often greatly elongate, capable of extending beyond the mantle cavity and can, apparently, transfer spermatophores directly to the female. terminal organ Wikipedia The long, muscular end of the male gonoduct that assists in transfering spermatophores to the female. Apparently, in species with a hectocotylus, the penis transfers spermatophores to the hectocotylus which in turn transfers them to the female. In species without a hectocotylus, the penis is often greatly elongate, capable of extending beyond the mantle cavity and can, apparently, transfer spermatophores directly to the female. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 terminal organ photocyte Cells in photophores that produce bioluminsecence. Cells in photophores that produce bioluminsecence. http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 photophore Organ that produces bioluminescence or &quot;living light.&quot; Photophores are often complex with various color filters, reflectors, light guides, lenses and chromatophores surrounding the site of light production, the photogenic region. The name of specific photophores often reflects their position on the animal. For example, the branchial photophore is at the base of the gill, the anal photophore lies at the side of the anus. light organ Organ that produces bioluminescence or &quot;living light.&quot; Photophores are often complex with various color filters, reflectors, light guides, lenses and chromatophores surrounding the site of light production, the photogenic region. The name of specific photophores often reflects their position on the animal. For example, the branchial photophore is at the base of the gill, the anal photophore lies at the side of the anus. http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 photophore location Photophores are found in nearly any location in some squid. However, certain locations are the most common. Photophores are found in nearly any location in some squid. However, certain locations are the most common. http://tolweb.org/accessory/Cephalopod_Photophore_Locations?acc_id=1981 photosensitive vesicle Non-image forming light-receptive organs located inside the cephalic cartilage in squid and on the stellate ganglion in octopods. They perform a variety of function. Non-image forming light-receptive organs located inside the cephalic cartilage in squid and on the stellate ganglion in octopods. They perform a variety of function. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 posterior buccal lobe A fused pair of lobes that are the anterior-most lobes of the supraesophageal mass of the brain of decapodiforms but is incorporated into the anterior portion of the supraesophageal mass behind the superior buccal lobes in most octopodiforms. inferior frontal lobe A fused pair of lobes that are the anterior-most lobes of the supraesophageal mass of the brain of decapodiforms but is incorporated into the anterior portion of the supraesophageal mass behind the superior buccal lobes in most octopodiforms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 inferior frontal lobe posterior lip gland Large glands surrounding the buccal mass just posterior to the lips. They are especially well-developed in stauroteuthid cirrates. Large glands surrounding the buccal mass just posterior to the lips. They are especially well-developed in stauroteuthid cirrates. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 posterior salivary gland Glands generally located just posterior to the head whose secretion travels via a duct into the buccal mass. Glands generally located just posterior to the head whose secretion travels via a duct into the buccal mass. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 primary conus protective membrane Membranes, generally supported by trabeculae, that form lateral borders along the oral surfaces of the arms and tentacular clubs in decapodiforms. A thin fold of integument along lateral angles of oral surface of arms and clubs lateral to suckers usually supported by trabeculae Membranes, generally supported by trabeculae, that form lateral borders along the oral surfaces of the arms and tentacular clubs in decapodiforms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A thin fold of integument along lateral angles of oral surface of arms and clubs lateral to suckers usually supported by trabeculae proximal and distal fields Some male opisthoteuthid cirrates have enlarged suckers located proximally on the arms (proximal fields) and/or distally on the arms (distal fields). Some male opisthoteuthid cirrates have enlarged suckers located proximally on the arms (proximal fields) and/or distally on the arms (distal fields). http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 pseudomorph A mass of ink and mucous that roughly has the dimensions of the cephalopod that secreted it. A mass of ink and mucous that roughly has the dimensions of the cephalopod that secreted it. http://tolweb.org/accessory/Cephalopod_Pseudomorph_Function?acc_id=1964 obsolete recent Refers to an organism that is living or has lived within the past 10,000 years or to object formed or events within the past 10,000 years. true Refers to an organism that is living or has lived within the past 10,000 years or to object formed or events within the past 10,000 years. http://tolweb.org/accessory/Geological_Timescale?acc_id=1987 photophore reflector Structures in a photophore that reflects bioluminscence. Structures in a photophore that reflects bioluminscence. http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 cephalopod radula A toothed ribbon that lies in the buccal mass and that aids in movement of food into the esophagus. A toothed ribbon that lies in the buccal mass and that aids in movement of food into the esophagus. http://tolweb.org/accessory/Cephalopod_Radula?acc_id=2035 renal appendage Structures that form the nephridium (=kidney). The renal appendages are outpocketings of the veins within the renal sac (primarily the venae cavae) that are covered with renal epithelium. The The renal sac communicates with the mantle cavity via the nephridial pore. kidney nephridium Structures that form the nephridium (=kidney). The renal appendages are outpocketings of the veins within the renal sac (primarily the venae cavae) that are covered with renal epithelium. The The renal sac communicates with the mantle cavity via the nephridial pore. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 rachis Central axis of the gladius, often thickened, that typically extends the full length of the gladius and sometimes called the median field. median field rhachis Central axis of the gladius, often thickened, that typically extends the full length of the gladius and sometimes called the median field. http://tolweb.org/accessory/Cephalopod_Gladius_Terminology?acc_id=1955 rostrum Portion of the gladius that extends posteriorly or posterodorsally from the conus. A distnct rostrum is found only in the Onychoteuthidae, Ancistrocheiridae, Lampadioteuthinae and Vampyroteuthidae. gladius Portion of the gladius that extends posteriorly or posterodorsally from the conus. A distnct rostrum is found only in the Onychoteuthidae, Ancistrocheiridae, Lampadioteuthinae and Vampyroteuthidae. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 saddle The region of the shell of cirrates connecting the lateral wings. The region of the shell of cirrates connecting the lateral wings. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 saddle-shaped The shape of the shell with an elongate central region (saddle proper) and broad wings found in cirroteuthid cirrates. The shape of the shell with an elongate central region (saddle proper) and broad wings found in cirroteuthid cirrates. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sagittate fin The most common fin-shape in squids with anterior free fin-lobes, a relatively sharp lateral angle and posterior elongation. See fin shapes. The most common fin-shape in squids with anterior free fin-lobes, a relatively sharp lateral angle and posterior elongation. See fin shapes. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 scales and tubercule The skin of the mantle contains scales, tubercules and warts in various species. todo - split The skin of the mantle contains scales, tubercules and warts in various species. http://tolweb.org/accessory/Cephalopod_Mantle_Armature?acc_id=1997 secondary conus A conical region at the posterior end of the gladius that is formed by an inrolling and fusion of the vanes. The ventral line of fusion is usually apparent. The secondary conus may be rather short or over half the gladius length. A conical region at the posterior end of the gladius that is formed by an inrolling and fusion of the vanes. The ventral line of fusion is usually apparent. The secondary conus may be rather short or over half the gladius length. http://tolweb.org/accessory/Cephalopod_Gladius_Terminology?acc_id=1955 secondary eyelid An eyelid that covers the ventral part of the cornea. secondary fold An eyelid that covers the ventral part of the cornea. http://tolweb.org/accessory/Cephalopod_Secondary_Eyelid?acc_id=1976 secondary fin A non-muscular fin-shaped structure found in some chiroteuthids, located posterior to the true or primary fin. The secondary fin may act as a buoyancy organ. A non-muscular fin-shaped structure found in some chiroteuthids, located posterior to the true or primary fin. The secondary fin may act as a buoyancy organ. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 secondary web Web that extends from each arm to the primary web in some incirrate octopods. Separation of the arm from the primary web seems to be associated with feeding and/or defense (bell and baloon balloon postures) but the function is not well understood. Web that extends from each arm to the primary web in some incirrate octopods. Separation of the arm from the primary web seems to be associated with feeding and/or defense (bell and baloon balloon postures) but the function is not well understood. http://tolweb.org/accessory/Cephalopod_Primary_and_Secondary_Webs?acc_id=1696 semelparous A reproductive strategy in which females spawn once then die. Sometimes called terminal or &quot;big-bang&quot; spawners. Many cephalopods are semelparous but many are not. A reproductive strategy in which females spawn once then die. Sometimes called terminal or &quot;big-bang&quot; spawners. Many cephalopods are semelparous but many are not. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 semicrescent plate (arm suckers) - A semicrescent plate on the proximal margin of the inner sucker arm rings of the loliginid squid <strong>Uroteuthis</strong>. (arm suckers) - A semicrescent plate on the proximal margin of the inner sucker arm rings of the loliginid squid <strong>Uroteuthis</strong>. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sepioid gill Gills of some cirrate octopods that look superficially like gills of sepioids. Gills of some cirrate octopods that look superficially like gills of sepioids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 shell sac The sac that secretes the shell in the Coleoidea. The sac is composed of ectodermal epithelium that invaginates during embryonic development to form an internal sac. The sac that secretes the shell in the Coleoidea. The sac is composed of ectodermal epithelium that invaginates during embryonic development to form an internal sac. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 side pocket Pockets formed by small membranous folds of integument lateral to the foveola in the funnel groove in ommastrephid and thysanoteuthid squids. Pockets formed by small membranous folds of integument lateral to the foveola in the funnel groove in ommastrephid and thysanoteuthid squids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 siphuncle The tube-like extension of the body wall, coelom and an external covering tube that penetrates the phragmocone and regulates gas exchange into the phragmocone chambers. http://en.wikipedia.org/wiki/Siphuncle The tube-like extension of the body wall, coelom and an external covering tube that penetrates the phragmocone and regulates gas exchange into the phragmocone chambers. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sperm mass The coiled rope of sperm that lies within the spermatophore or spermatangia. rope of sperm todo The coiled rope of sperm that lies within the spermatophore or spermatangia. http://tolweb.org/accessory/Cephalopod_Spermatophore_Terminology?acc_id=1972 spermatangia Discharged spermatophore which consists of the sperm mass surrounded by a membrane (the inner tunic) with the cement gland at one end. Discharged spermatophore which consists of the sperm mass surrounded by a membrane (the inner tunic) with the cement gland at one end. http://tolweb.org/accessory/Cephalopod_Spermatophore_Terminology?acc_id=1972 cephalopod spermathecum Specialized sperm-storage structures found in the skin of some female decapodiforms or as pockets of the oviducal gland in octopods. Specialized sperm-storage structures found in the skin of some female decapodiforms or as pockets of the oviducal gland in octopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 spermatophore A packet of sperm that is formed by the male and passed to the female during mating. In most coleoids, this packet is very complex and contains a &quot;rope&quot; of sperm (= sperm mass), an ejaculatory apparatus and a cement body. Except for the sperm, the entire structure is non-cellular and consists of a complex architecture of secreted material. todo - create subclass for undischarged form A packet of sperm that is formed by the male and passed to the female during mating. In most coleoids, this packet is very complex and contains a &quot;rope&quot; of sperm (= sperm mass), an ejaculatory apparatus and a cement body. Except for the sperm, the entire structure is non-cellular and consists of a complex architecture of secreted material. http://tolweb.org/accessory/Cephalopod_Spermatophore_Terminology?acc_id=1972 stalk of tentacle Region of the tentacle proximal to the club. Region of the tentacle proximal to the club. http://tolweb.org/accessory/Cephalopod_Tentacle_Terminology?acc_id=1962 stalked eye cephalopod statocyst A sense-organ that detects gravity, angular accelerations and low-frequency sound. The statocyst is embedded within the cephalic cartlage and contains the statolith. A sense-organ that detects gravity, angular accelerations and low-frequency sound. The statocyst is embedded within the cephalic cartlage and contains the statolith. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 stellate ganglion Major ganglion of the peripherial nervous system of neocoleoid cephalopods that controls nerves to the mantle muscles. Major ganglion of the peripherial nervous system of neocoleoid cephalopods that controls nerves to the mantle muscles. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod stomach A muscular organ of the digestive system where primary digestion occurs. The stomach is generally lined with cuticular ridges to aid in grinding food, and is supplied with digestive enzymes from the digestive gland. The stomach may be greatly expandable in size and serve as a storage area until food can be fully processed. A muscular organ of the digestive system where primary digestion occurs. The stomach is generally lined with cuticular ridges to aid in grinding food, and is supplied with digestive enzymes from the digestive gland. The stomach may be greatly expandable in size and serve as a storage area until food can be fully processed. http://tolweb.org/accessory/Cephalopod_Digestive_System?acc_id=1993 stylet Remnant of the molluscan shell in incirrate octopods. It is generally a slender, pointed rod tightly surrounded by the shell sac and buried in the mantle muscle on either side of the mantle. Remnant of the molluscan shell in incirrate octopods. It is generally a slender, pointed rod tightly surrounded by the shell sac and buried in the mantle muscle on either side of the mantle. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 subadult stage Stage at which all of the characters that typically define the species are present. The subadult stage follows the Juvenile Stage and precedes the Adult Stage. A subadult stage is defined in cephalopods since the adult phase is frequently abbreviated. Stage at which all of the characters that typically define the species are present. The subadult stage follows the Juvenile Stage and precedes the Adult Stage. A subadult stage is defined in cephalopods since the adult phase is frequently abbreviated. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sucker Muscular suction-cup on the arms and tentacles and occasionally on the buccal supports. The sucker consists of a cup-shaped portion, the acetabulum, and a flat, distal ring, the infundibulum, that contacts the substrate. Suckers of decapodiforms contain horny rings that often bear teeth or sometimes form hooks. Muscular suction-cup on the arms and tentacles and occasionally on the buccal supports. The sucker consists of a cup-shaped portion, the acetabulum, and a flat, distal ring, the infundibulum, that contacts the substrate. Suckers of decapodiforms contain horny rings that often bear teeth or sometimes form hooks. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sucker series The longitudinal rows of suckers on the arms or tentacles of decapodiforms. Series (= longitudinal rows) contrasts with rows (= transverse rows) in describing sucker arrangement. In octopodiforms, sucker &quot;series&quot; is generally synonymous with &quot;rows&quot;, the &quot;longitudinal&quot; component of the term is implied. Suckers can form distinct, regular series or indistinct, irregular series. The longitudinal rows of suckers on the arms or tentacles of decapodiforms. Series (= longitudinal rows) contrasts with rows (= transverse rows) in describing sucker arrangement. In octopodiforms, sucker &quot;series&quot; is generally synonymous with &quot;rows&quot;, the &quot;longitudinal&quot; component of the term is implied. Suckers can form distinct, regular series or indistinct, irregular series. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sucker stalk The structure between the sucker and the arm. In octopods it is a cyclindrical structure about the same width as the sucker and in decapodiforms it is constricted into a conical pillar. pedicel The structure between the sucker and the arm. In octopods it is a cyclindrical structure about the same width as the sucker and in decapodiforms it is constricted into a conical pillar. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 sucker teeth Teeth on the inner horny sucker rings of some decapodiforms. Teeth on the inner horny sucker rings of some decapodiforms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 superior buccal lobes Lobes of the central nervous system that sit atop the esophagus where the latter enters the buccal mass in decapodiforms but are the anterior-most lobes of the supraesophageal mass of the brain of octopodiforms. Lobes of the central nervous system that sit atop the esophagus where the latter enters the buccal mass in decapodiforms but are the anterior-most lobes of the supraesophageal mass of the brain of octopodiforms. http://tolweb.org/accessory/Cephalopod_Brain_Terminology?acc_id=1944 superocular cirri Muscular structures that can be raised or lowered above the eyes of octopodids but are apparent even when lowered. Muscular structures that can be raised or lowered above the eyes of octopodids but are apparent even when lowered. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 suprabrachial commissure Nerve commissure that passes dorsal to the esophagus in most octopodiforms and connects the right and left anterior subesophageal masses of the brain. todo - mireot Nerve commissure that passes dorsal to the esophagus in most octopodiforms and connects the right and left anterior subesophageal masses of the brain. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod swimbladder Gas-filled structure found in the dorsal region of the pelagic octopods, <em><strong>Ocythoe</strong>,</em> <em><strong>Tremoctopus</strong></em> and <em><strong>Haliphron</strong></em>. gas bladder swim bladder Gas-filled structure found in the dorsal region of the pelagic octopods, <em><strong>Ocythoe</strong>,</em> <em><strong>Tremoctopus</strong></em> and <em><strong>Haliphron</strong></em>. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 tentacle Modified 4th pair of appendages in decapodiforms that are capable of considerable extension and contraction. Although the tentacles are derived evolutionarily from the 4th pair of arms, the term, "arms IV"; is reserved for the subsequent arm pair, the ventral arms, which are evolutionarily the 5th pair of arms. Each tentacle consist of a proximal stalk usually devoid of suckers, and a distal club armed with suckers and occasionally hooks. arm limb Elongate 4th circumoral appendage of cuttlefish and squids used to capture prey. Divided into proximal stalk and distal club. Modified 4th pair of appendages in decapodiforms that are capable of considerable extension and contraction. Although the tentacles are derived evolutionarily from the 4th pair of arms, the term, "arms IV"; is reserved for the subsequent arm pair, the ventral arms, which are evolutionarily the 5th pair of arms. Each tentacle consist of a proximal stalk usually devoid of suckers, and a distal club armed with suckers and occasionally hooks. Wikipedia:Cephalopod_limb http://tolweb.org/accessory/Composite_Squid?acc_id=1936 Wikipedia:Cephalopod_limb limb http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Elongate 4th circumoral appendage of cuttlefish and squids used to capture prey. Divided into proximal stalk and distal club. Wikipedia:Cephalopod_limb arm tentacle absence Tentacles can be absent because the species lacks tentacles, they are accidentally lost during capture, or they are naturally lost at sexual maturity. Tentacles can be absent because the species lacks tentacles, they are accidentally lost during capture, or they are naturally lost at sexual maturity. http://tolweb.org/accessory/Tentacle_Absence_in_Cephalopods?acc_id=2046 tentacle pad Poorly understood and complex pad-like photophores that are found on the tentacular stalks of some chiroteuthidids. Poorly understood and complex pad-like photophores that are found on the tentacular stalks of some chiroteuthidids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 tentacle pocket A pocket that surrounds the base of each tentacle and provides some room for tentacle retraction or contraction. tentacular pocket A pocket that surrounds the base of each tentacle and provides some room for tentacle retraction or contraction. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 tentacle thickness . . http://tolweb.org/accessory/Cephalopod_Tentacle_Thickness?acc_id=2016 terminal fin Fins that extend to the posterior end of the body. Fins that extend to the posterior end of the body. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 terminal pad A circlet or disc of small suckers at the tip of the tentacular club. The pad of one club can adhere to the pad of the other club which holds the clubs together at their tips. terminal pad of tentacular club A circlet or disc of small suckers at the tip of the tentacular club. The pad of one club can adhere to the pad of the other club which holds the clubs together at their tips. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 trabecula Muscular cones or flaps that support the protective membranes on the arms and tentacular clubs in decapodiforms. These are thought to be homologous with the cirri of the octopodiformes. trabeculae Muscular cones or flaps that support the protective membranes on the arms and tentacular clubs in decapodiforms. These are thought to be homologous with the cirri of the octopodiformes. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 tragus Particular inward projecting knob in the funnel locking-apparatus of some squids. It is the knob found on the medial surface of the central depression. Particular inward projecting knob in the funnel locking-apparatus of some squids. It is the knob found on the medial surface of the central depression. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 truncate tooth Teeth on the inner horny of rings of decapodiform suckers that do not terminate in a point but rather a broad, flat tip. truncate teeth Teeth on the inner horny of rings of decapodiform suckers that do not terminate in a point but rather a broad, flat tip. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod tubercule Structures with one or more cusps just beneath the surface epithelium. Tubercules may be cartilagenous, fibrous or of other structure and may be confined to the integument or extend deeper. Structures with one or more cusps just beneath the surface epithelium. Tubercules may be cartilagenous, fibrous or of other structure and may be confined to the integument or extend deeper. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 gladius vane Broad, lateral portions of the gladius that arise from the rhachis. lateral field wing vane Broad, lateral portions of the gladius that arise from the rhachis. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 ventral eyelid An eyelid that can cover much of the cornea from a ventral position in many sepioids. An eyelid that can cover much of the cornea from a ventral position in many sepioids. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 ventral shield The optically modified ventral mantle of heteroteuthins that usually projects anteriorly beneath the head. The optically modified ventral mantle of heteroteuthins that usually projects anteriorly beneath the head. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 vesicular tissue Tissues found in some decapodiforms that contain numberous vesicles filled with water that usually contains ammonium chloride. The vesicles may be concentrated in certain structures (e.g., the ventral arms) or scattered throughout the animal. The light solution in the vesicles provide buoyant lift. Tissues found in some decapodiforms that contain numberous vesicles filled with water that usually contains ammonium chloride. The vesicles may be concentrated in certain structures (e.g., the ventral arms) or scattered throughout the animal. The light solution in the vesicles provide buoyant lift. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 visceral nucleus A concentration of digestive organs that sometimes includes reproductive organs as well. In cranchiid squids the visceral nucleus is especially distinct: the small, spindle-shaped digestive gland contains the ink sac and is adjacent to the stomach and caecum and associated structures (eg, digestive gland duct appendages, renal appendages). In most cephalopods the concentration of these structures is not so extensive but whenever a concentration exists, a visceral nucleus is present. BTO:0001448 visceral hump A concentration of digestive organs that sometimes includes reproductive organs as well. In cranchiid squids the visceral nucleus is especially distinct: the small, spindle-shaped digestive gland contains the ink sac and is adjacent to the stomach and caecum and associated structures (eg, digestive gland duct appendages, renal appendages). In most cephalopods the concentration of these structures is not so extensive but whenever a concentration exists, a visceral nucleus is present. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 BTO:0001448 visceral hump visceral sac The body posterior to the head surrounded by the mantle. The body wall in this region that incases the viscera is usually rather thin-walled, hence the name &quot;visceral sac.&quot; The visceral sac is also called the &quot;visceral dome.&quot; visceral dome The body posterior to the head surrounded by the mantle. The body wall in this region that incases the viscera is usually rather thin-walled, hence the name &quot;visceral sac.&quot; The visceral sac is also called the &quot;visceral dome.&quot; http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 visceropericardial coelom The largest coelom in cephalopods except in the Octopoda where it is greatly reduced. Geneally it encloses the gonad, and partially the stomach, caecum and ventricle among other structures. Eggs and sperm are released into the coelom and picked up by the gonoducts. The visceropercardial coelom also communicates with the other cephalopod coelom, the nephridial coelom, and the mantle cavity via a pair of ducts that open at the base of the nephridial papillae. The largest coelom in cephalopods except in the Octopoda where it is greatly reduced. Geneally it encloses the gonad, and partially the stomach, caecum and ventricle among other structures. Eggs and sperm are released into the coelom and picked up by the gonoducts. The visceropercardial coelom also communicates with the other cephalopod coelom, the nephridial coelom, and the mantle cavity via a pair of ducts that open at the base of the nephridial papillae. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 water pore One or two pairs of pores in the integument of the head that communicate with extensive spaces at the bases of arms on some pelagic argonautoid octopods. One or two pairs of pores in the integument of the head that communicate with extensive spaces at the bases of arms on some pelagic argonautoid octopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 web Muscular membrane that extends between the arms of many octopodiforms but is reduced or absent in most decapodiforms. Muscular membrane that extends between the arms of many octopodiforms but is reduced or absent in most decapodiforms. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 web nodule Muscular rods embedded in the web of some cirrate octopods. Muscular rods embedded in the web of some cirrate octopods. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 white body Amorphous organ near the optic lobes that forms white blood cells. TODO: immune system Amorphous organ near the optic lobes that forms white blood cells. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 wing (shell) - Lateral limbs of the shell of cirrate octopods (shell) - Lateral limbs of the shell of cirrate octopods http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 leucophore Reflective cells in the skin of some cephalopods containing numerous refractive granules on stalks that together produce a scattered reflection of ambient light. TODO: cells Reflective cells in the skin of some cephalopods containing numerous refractive granules on stalks that together produce a scattered reflection of ambient light. http://tolweb.org/notes/?note_id=2646 ink primary oviduct Female gonoduct. The oviduct conducts eggs from the visceropericardial coelom, that holds the ovary, to the exterior and is often used to store eggs. In some argonautoid octopods eggs are fertilized and undergo either partial (Argonauta ) or complete (Ocythoe) embryonic development within the oviduct. Female gonoduct. The oviduct conducts eggs from the visceropericardial coelom, that holds the ovary, to the exterior and is often used to store eggs. In some argonautoid octopods eggs are fertilized and undergo either partial (Argonauta ) or complete (Ocythoe) embryonic development within the oviduct. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 cephalopod ovary photosensitive vesicle of cephalic cartilage squid photosensitive vesicle of stellate ganglion octopod obsolete brachial photophore true tentacular club Terminal, usually expanded, grasping portion of tentacle that is armed with suckers or, in some groups, with suckers and hooks. Terminal, usually expanded, grasping portion of tentacle that is armed with suckers or, in some groups, with suckers and hooks. Terminal, usually expanded, grasping portion of tentacle that is armed with suckers or, in some groups, with suckers and hooks. optic nerve bundle Large nerve bundles that run from the optic lobe to the eye. optic stalk todo - mireot Large nerve bundles that run from the optic lobe to the eye. http://tolweb.org/accessory/Cephalopoda_Glossary?acc_id=587 lobe of brain lobe of nervous system buccal lobe subesophageal mass of brain middle subesophageal mass of brain posterior subesophageal mass of brain supraesophageal mass of brain cephalopod trait 2011-01-30T08:15:10Z cjm basal lobe of brain Higher motor centers, located (for example) in the basal lobe (b.l.) control complex behaviors that involve the animal's whole body. superior frontal lobe system vertical frontal lobe system VL vertical lobe anal photophore photophore at side of anus http://tolweb.org/accessory/Cephalopod_Photophore_Terminology?acc_id=2015 photophore at side of anus collection of suckers transverse row of suckers circumoral appendage segment TODO - mireot from uberon circumoral appendage arm crown head appendage Any of the numerous arms or tentacles that arise from the head and surround the mouth of a cephalopod http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Any of the numerous arms or tentacles that arise from the head and surround the mouth of a cephalopod cephalopod size http://en.wikipedia.org/wiki/Cephalopod_size buccal membrane support buccal support longitudinal muscular rod fused with buccal membrane bullet-shaped mantle posteriorly blunt rather than broad body (mantle) common in larval cephalopods http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf posteriorly blunt rather than broad body (mantle) common in larval cephalopods extrategumental chromatophore A larval chromatophore located deep beneather the integument http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A larval chromatophore located deep beneather the integument chromatophore field Collection of chromatophores that produces species-specific patterns in discrete regions http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Collection of chromatophores that produces species-specific patterns in discrete regions tegumental chromatophore Chromatophores located in or near surface integument http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Chromatophores located in or near surface integument cirrus body cirrus A fleshy protruberance of skin that can be erected as a papilla, usually dorsal to the eyes http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A fleshy protruberance of skin that can be erected as a papilla, usually dorsal to the eyes phragmacone found in fossil teuthoids The phragmocone is the chambered portion of the shell of a cephalopod. It is divided by septa into camerae. http://en.wikipedia.org/wiki/Phragmocone The phragmocone is the chambered portion of the shell of a cephalopod. It is divided by septa into camerae. http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf found in fossil teuthoids http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf corneal membrane A thin turgid transparent membrane that covers the eyes of myopsid and sepioid cephalopods http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A thin turgid transparent membrane that covers the eyes of myopsid and sepioid cephalopods gill length Koelicker organ Minute bristle-like structure that covers the body of planktonic octopod larvae lanceola Expanded portion of gladius vane http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Expanded portion of gladius vane macula Irregular spot capable of expanding or retracting nuchal fold One of a series of longitudinal folds or pleats of skin in the nuchal region http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf One of a series of longitudinal folds or pleats of skin in the nuchal region ocellus false eye spot cephalopod sperm receptacle A bulbous structure found in buccal region of femles of some cephalopods for retention of viable sperm until required for fertilization http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf A bulbous structure found in buccal region of femles of some cephalopods for retention of viable sperm until required for fertilization spermatophore pad inner sucker ring outer sucker ring tail Posterior extension of gladius and mantle epithelium. Frequently elongate. http://www.sil.si.edu/smithsoniancontributions/zoology/pdf_hi/sctz-0513.pdf Posterior extension of gladius and mantle epithelium. Frequently elongate. branchial heart appendage pericardial gland spot dorsal beak upper mandible ventral beak lower mandible septum arm I dorsal arm arm II arm III external shell Nautiluses are the only living cepholopod with an external shell throughout their life cycle ftp://ftp.fao.org/docrep/fao/009/w7192e/w7192e02.pdf Nautiluses are the only living cepholopod with an external shell throughout their life cycle spermatophore groove A channel or groove running along the length of the hectocotylus ftp://ftp.fao.org/docrep/fao/009/w7192e/w7192e02.pdf A channel or groove running along the length of the hectocotylus shield dorsal shield hard calcareous dorsal surface of the cuttlebone ftp://ftp.fao.org/docrep/fao/009/w7192e/w7192e02.pdf hard calcareous dorsal surface of the cuttlebone epirenal body glandular structures of unknown function which lie on either side of the renal papillae ftp://ftp.fao.org/docrep/fao/009/w7192e/w7192e02.pdf glandular structures of unknown function which lie on either side of the renal papillae renal sac kidney nephridium tooth of sucker ring umbilicus the central core of the chambered Nautilus shell, representing the juvenile shell with its initial coils ftp://ftp.fao.org/docrep/fao/009/w7192e/w7192e02.pdf the central core of the chambered Nautilus shell, representing the juvenile shell with its initial coils web depth beak length hood length lower rostral length LRL upper rostral length URL spadix organ shell chamber Camerae (singular camera) are the spaces or chambers enclosed between two adjacent septa in the phragmocone of a nautiloid or ammonoid cephalopod. These can be seen in cross-sections of a nautilus shell and in the polished cross-sections of ammonites. In life these chambers are filled with gas, mediated by the siphuncle, and used to control buoyancy. phragmacone chamber http://en.wikipedia.org/wiki/Camera_%28cephalopod%29 Camerae (singular camera) are the spaces or chambers enclosed between two adjacent septa in the phragmocone of a nautiloid or ammonoid cephalopod. These can be seen in cross-sections of a nautilus shell and in the polished cross-sections of ammonites. In life these chambers are filled with gas, mediated by the siphuncle, and used to control buoyancy. shell septum orthocone An orthocone is a usually long straight shell of a nautiloid cephalopod. http://en.wikipedia.org/wiki/Orthocone An orthocone is a usually long straight shell of a nautiloid cephalopod. internal shell shell aperture An opening in a mollusc shell http://en.wikipedia.org/wiki/Aperture_%28mollusc%29 An opening in a mollusc shell shell apex the tip of the mollusc shell of a gastropod, scaphopod, or cephalopod mollusc. n orthocone cephalopods, the pointed end of the shell is called the apex, and shell growth is away from the apex and toward the aperture. The first chamber of the apex is sometimes called the protoconch. http://en.wikipedia.org/wiki/Apex_%28mollusc%29 the tip of the mollusc shell of a gastropod, scaphopod, or cephalopod mollusc. funnel component of funnel-mantle locking apparatus mantle component of funnel-mantle locking apparatus spermatophore gland Ammonoidea ammonites http://purl.obolibrary.org/obo/NCBITaxon_subclass Goniatitida http://purl.obolibrary.org/obo/NCBITaxon_order Ceratitida http://purl.obolibrary.org/obo/NCBITaxon_order Ammonitida http://purl.obolibrary.org/obo/NCBITaxon_order reproductive trait appendage trait habitat trait In future, this class will be connected to ENVO shell trait integument trait fin trait head trait whole organism trait gill trait iteroparous characterized by multiple reproductive cycles over the course of its lifetime Nautiluses are the only cephalopods to present iteroparity organ of Hoeven pyroform sac An organ on the left side of the reproductive tract occupying the analagous position to the genital duct in males and females. Found in Nautilus. pyroform gland sexual size dimorphism Tremoctopus exhibit an extreme degree of sexual dimorphism. Females may reach 2 m in length, whereas the tiny males are at most a few centimeters long. sexual dimorphism buccal ganglion epithelial layer of gladius gladius epithelium pen epithelium epithelial layer of pen marginal epithelial layer of the gladius marginal epithelial layer on the pen ventral epithelial layer of gladius ventral epithelial layer on the pen iridophore leucophore biological_process Any process specifically pertinent to the functioning of integrated living units: cells, tissues, organs, and organisms. A process is a collection of molecular events with a defined beginning and end. Any process specifically pertinent to the functioning of integrated living units: cells, tissues, organs, and organisms. A process is a collection of molecular events with a defined beginning and end. GOC:go_curators GOC:isa_complete bioluminescence The production of light by certain enzyme-catalyzed reactions in cells. Wikipedia:Bioluminescence ISBN:0198506732 The production of light by certain enzyme-catalyzed reactions in cells. locomotion Self-propelled movement of a cell or organism from one location to another. GOC:dgh Self-propelled movement of a cell or organism from one location to another. flight Self-propelled movement of an organism from one location to another through the air, usually by means of active wing movement. GOC:dph Self-propelled movement of an organism from one location to another through the air, usually by means of active wing movement.